Beetles battling for mating rights, a bear stalking through the rainforest and a damselfly perched in front of a waterfall ...

While the Top 50 remains densely populated by gastropubs from London and the South East, the North of England claimed 15 of the 50 top spots in this year's list, three of which belong to Cumbrian ...

The stunning winners are a diverse range of intimate portraits, infrared and iPhone photography, and striking black-and-white images ...